Document Type : Final File


1 Department of Ecology, Institute of Science, High Technology and Environmental Science, Graduate University of Advanced Technology, Kerman, Islamic Republic of Iran

2 Department of Geology, Faculty of Sciences, University of Isfahan, Isfahan, Islamic Republic of Iran


The aim of this study is to bridge a gap in the existence of Trilobites in the marine Permian successions of Iran by exploring a stratigraphic section of the Permian in Jamal Formation (Dizlu section).  Based on comprehensive field studies and laboratory investigations three Trilobite specimens were identified in the study area including Pseudophillipsia (Carniphillipsia) dizluensis sp. nov., Pseudophillipsia (Carniphillipsia) sagittalis and Pseudophillipsia (Carniphillipsia) sp.. A new occurrence of trilobites collected from the Permian (Wordian) Jamal Formation at a section at Dizlu, Isfahan Province, Iran, is documented.   They belong to a new species of Pseudophillipsia (Carniphillipsia), which is described herein, and can be attributed to the a group of species centered on P.(C). paffenholzi (Weber). It is based on 30 specimens from a single horizon, most of which are enrolled. Co-occurrence of Pseudophillipsia (Carniphillipsia) sp. A. and stenohalin groups such as brachiopods, bryozoans, and crinoids are indicating that this species were preferred shallow and carbonate environments. This is the first time that the existences of these specimens are reported in Dizlu section.  


Main Subjects

1. Taraz H., Golshani F., Nakazawa K., Shimizu D., Bando Y., Ishii K. I., Murata M., Okimura Y., Sakagami S. & Nakamura K. The Permian and the Lower Triassic systems in Abadeh region, central Iran.  (1981).
2.  Lerosey-Aubril R. The Late Palaeozoic trilobites of Iran and Armenia and their palaeogeographical significance. Geological Magazine, 149:(06): 1023-1045 (2012)
3.  Lerosey-Aubril R. & Angiolini L. Permian trilobites from Antalya Province, Turkey, and enrollment in Late Palaeozoic trilobites. Turkish Journal of Earth Sciences, 18:(3): 427-448 (2009).
4.  Fortey R. A. & Heward A. P. A new, morphologically diverse Permian trilobite fauna from Oman. Acta Palaeontol Pol., 60:(1): 201-216 (2015).
5.  Mistiaen B., Gholamalian H., Gourvennec R., Plusquellec Y., Bigey F., Brice D., Feist M., Feist R., Pour M. G. & Kebria-Ee M. Preliminary data on the Upper Devonian (Frasnian, Famennian) and Permian fauna and flora from the Chahriseh area (Esfahan Province, central Iran). Annales-Societe Geologique Du Nord, 8:(2): 93-102 (2000).
6.  Ernst A., Senowbari-Daryan B. & Hamedani A. Middle Permian Bryozoa from the Lakaftari area, northeast of Esfahan (central Iran). Geodiversitas, 28:(4): 543-590 (2006).
7.  Ameri H. Peri-Gondwana Late Early–Middle Cambrian trilobites from the Kuhbanan Formation in Dahu section, Kerman Province, Iran. Arabian Journal of Geosciences: 1-12 (2013).
8.  Stampfli G. & Borel G. A plate tectonic model for the Paleozoic and Mesozoic constrained by dynamic plate boundaries and restored synthetic oceanic isochrons. Earth and Planetary Science Letters, 196:(1): 17-33 (2002).
9.  Kozur H. Correlation of the continental uppermost Permian and Lower Triassic of the Germanic Basin with the marine scale in the light of new data from China and Iran. Albertiana, 33: 48-51 (2005).
10. Heydari E., Arzani N., Safaei M. & Hassanzadeh J. Ocean's response to a changing climate: Clues from variations in carbonate mineralogy across the Permian–Triassic boundary of the Shareza Section, Iran. Global Planet. Change, 105: 79-90 (2013).
11. Kozur H. Biostratigraphy and event stratigraphy in Iran around the Permian–Triassic Boundary (PTB): implications for the causes of the PTB biotic crisis. Global Planet. Change, 55:(1): 155-176 (2007).
12. Arefifard S. & Isaacson P. E. Permian Sequence stratigraphy in east-central Iran: Microplate records of Peri-Tethyan and Peri-Gondwanan events. Stratigraphy, 8:(1): 61-83 (2011).
13. Richoz S., Krystyn L., Baud A., Brandner R., Horacek M. & Mohtat-Aghai P. Permian–Triassic boundary interval in the Middle East (Iran and N. Oman): Progressive environmental change from detailed carbonate carbon isotope marine curve and sedimentary evolution. J. Asian Earth Sci. 39:(4): 236-253 (2010).
14.  Liu X. C., Wang W., Shen S. Z., Gorgij M. N., Ye F. C., Zhang Y. C., Furuyama S., Kano A. & Chen X. Z. Late Guadalupian to Lopingian (Permian) carbon and strontium isotopic chemostratigraphy in the Abadeh section, central Iran. Gondwana Research., 24:(1): 222-232 (2013).
15.  Shen S. Z. & Mei S. L. Lopingian (Late Permian) high‐resolution conodont biostratigraphy in Iran with comparison to South China zonation. Geological Journal., 45:(2‐3): 135-161 (2010).
16.  Ghaderi A., Leda L., Schobben M., Korn D. & Ashouri A. High-resolution stratigraphy of the Changhsingian (Late Permian) successions of NW Iran and the Transcaucasus based on lithological features, conodonts and ammonoids. - Mitteilungen aus dem Museum für Naturkunde in Berlin. Fossil Record., 17:(1): 41 (2014).
17.  Leda L., Korn D., Ghaderi A., Hairapetian V., Struck U. & Reimold W. U. Lithostratigraphy and carbonate microfacies across the Permian–Triassic boundary near Julfa (NW Iran) and in the Baghuk Mountains (Central Iran). Facies., 60:(1): 295-325 (2014).
18.  Leven E. J. & Gorgij M. Fusulinids and stratigraphy of the Carboniferous and Permian in Iran. Stratigraphy and Geological Correlation., 19:(7): 687-776 (2011).
19.  Zakharov Y. D. & Abnavi N. M. The ammonoid recovery after the end-Permian mass extinction: evidence from the Iran-Transcaucasia area, Siberia, Primorye, and Kazakhstan. Acta Palaeontol. Pol., 58:(1): 127-147 (2013).
20.  Davydov V. I. & Arefifard S. Middle Permian (Guadalupian) fusulinid taxonomy and biostratigraphy of the mid-latitude Dalan Basin, Zagros, Iran and their applications in paleoclimate dynamics and paleogeography. GeoArabia, Journal of the Middle East Petroleum Geosciences., 18:(2): 17-62 (2013).
21.  Moghadam H. S., Li X. H., Ling X. X., Stern R. J., Khedr M. Z., Chiaradia M., Ghorbani G., Arai S. & Tamura A. Devonian to Permian evolution of the Paleo-Tethys Ocean: New evidence from U–Pb zircon dating and Sr–Nd–Pb isotopes of the Darrehanjir–Mashhad “ophiolites”, NE Iran. Gondwana Research., 28:(2): 781-799 (2015).
22. Ameri H. Peri-Gondwana Late Early–Middle Cambrian trilobites from the Kuhbanan Formation in Dahu section, Kerman Province, Iran. Arabian Journal of Geosciences., 8:(3): 1467-1478 (2015).
23. Hampe O., Hairapetian V., Dorka M., Witzmann F., Akbari A. M. & Korn D. A first late Permian fish fauna from Baghuk Mountain (Neo-Tethyan shelf, central Iran). Bull. Geosci., 88:(1): 1-20 (2013).
24. Nejad M. E., Vachard D., Siabeghodsy A. A. & Abbasi S. Middle-Late Permian (Murgabian-Djulfian) foraminifers of the northern Maku area (western Azerbaijan, Iran). Palaeontologia Electronica., 18:(1): 1-63 (2015).